Tim Guilford

Professor of Animal Behaviour

Tutorial Fellow in Zoology, Merton College


Although I still maintain an interest in the evolution of animal signals, especially warning signals, which was the subject of my early research, I now concentrate on trying to unravel the mechanisms by which animals map the environments through which they travel. What cues do birds use to navigate home across familiar areas? What do their maps look like? How versatile is the spatial knowledge they store? The questions range from cognitive to ecological, homing pigeons are the principal model, and the techniques involve field experiments using miniature GPS loggers, on-board cameras, and analytical approaches borrowed from computational engineering. Most recently I have been extending my research group’s work to trying to understand how long distance wandering seabirds (many of which are of vulnerable conservation status) migrate, map and navigate across the open oceans.

Contact Details

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Zoology Research and Administration Building
11a Mansfield Road
           Oxford OX1 3SZ

tim guilford



  • Young frigatebirds learn how to compensate for wind-drift

    Wynn, J, COLLET, J, Prudor, A, Corbeau, A, Padget, O, GUILFORD, T, Weimerskirch, H
  • Short-term behavioural impact contrasts with long-term fitness consequences of biologging in a long-lived seabird

    Gillies, N, FAYET, A, Padget, O, Syposz, M, Wynn, J, Bond, S, Evry, J, Kirk, H, Shoji, A, Dean, B, Freeman, R, GUILFORD, T
  • Natal imprinting to the Earth’s magnetic field in a pelagic seabird

    GUILFORD, T, Wynn, J, Padget, O, Mouritsen, H, Perrins, C
  • Manx shearwater (Puffinus puffinus) rafting behaviour revealed by GPS tracking and behavioural observations

    Richards, C, Padget, O, Guilford, T, Bates, AE
  • Patterns of at-sea behaviour at a hybrid zone between two threatened seabirds.

    Austin, RE, Wynn, RB, Votier, SC, Trueman, C, McMinn, M, Rodríguez, A, Suberg, L, Maurice, L, Newton, J, Genovart, M, Péron, C, Grémillet, D
    et al
  • Shearwaters know the direction and distance home but fail to encode intervening obstacles after free-ranging foraging trips.

    Padget, O, Stanley, G, Willis, JK, Fayet, AL, Bond, S, Maurice, L, Shoji, A, Dean, B, Kirk, H, Juarez-Martinez, I, Freeman, R, Bolton, M
    et al
  • Light pollution causes object collisions during local nocturnal manoeuvring flight by adult Manx shearwaters (Puffinus puffinus)

    GUILFORD, T, Padget, O, Bond, S, Syposz, M
  • The shearwater's world

  • Spatial scales of marine conservation management for breeding seabirds

    Oppel, S, Bolton, M, Carneiro, APB, Dias, MP, Green, JA, Masello, JF, Phillips, RA, Owen, E, Quillfeldt, P, Beard, A, Bertrand, S, Blackburn, J
    et al
  • Predicting animal behaviour using deep learning: GPS data alone accurately predict diving in seabirds

    Browning, E, Bolton, M, Owen, E, Shoji, A, Guilford, T, Freeman, R
  • More